The 2016 National Geographic Nature Photographer of the Year contest announced the winners in four categories: landscape, environmental issues, action and animal portraits. The following entries are the top prizes for each category and the overall grand prize.

"During the sardine migration along the Wild Coast of South Africa, millions of sardines are preyed upon by marine predators such as dolphins, marine birds, sharks, whales, penguins, sailfishes, and sea lions."
Greg Lecoeur/2016 National Geographic Nature Photographer of the Year